Cambridge University Press
978-0-521-85726-0 - Zooarchaeology - Second Edition - by Elizabeth J. Reitz
Frontmatter/Prelims



Zooarchaeology, Second Edition



This book serves as an introductory text for students interested in the identification and the analysis of animal remains from archaeological sites. The emphasis is on animals whose remains inform us about the relationship between humans and their natural and social environments, especially site-formation processes, subsistence strategies, the processes of domestication, and paleoenvironments. Examining studies from all over the world, from the Pleistocene period up to the present, this volume is organized in a way that is parallel to a faunal study, beginning with background information, bias in a faunal assemblage, and basic zooarchaeological methods. A zooarchaeological Hypothetical Collection illustrates the fundamental methods for the collection of primary and secondary data that are applicable to zooarchaeology practice.

   This revised edition reflects developments in zooarchaeology that have occurred during the past decade. It includes new sections on enamel ultrastructure and incremental analysis, stable isotopes and trace elements, ancient genetics and enzymes, environmental reconstruction, people as agents of environmental change, applications of zooarchaeology in animal conversation and heritage management, and a discussion of issues pertaining to the curation of zooarchaeological materials.

ELIZABETH J. REITZ is Professor of Anthropology at the Georgia Museum of Natural History, University of Georgia. Her work is based on the identification and interpretation of animal remains from coastal archaeological sites, particularly in South America, the Caribbean, and the southeastern United States. She is the co-author and co-editor of several volumes, as well as the author of more than 150 articles and book chapters.

ELIZABETH S. WING is Curator Emeritus at the Florida Museum of Natural History, University of Florida. The co-author of two books and author of many scholarly articles, she received the Fryxell Award from the Society for American Archaeology in 1996 for distinguished contributions to archaeology through interdisciplinary research. In 2006, Dr. Wing was elected to the National Academy of Sciences and received the President’s Medal from the University of Florida.





Cambridge Manuals in Archaeology

General Editor
Graeme Barker, University of Cambridge

Advisory Editors
Elizabeth Slater, University of Liverpool
Peter Bogucki, Princeton University


Cambridge Manuals in Archaeology is a series of reference handbooks designed for an international audience of upper-level undergraduate and graduate students, and professional archaeologists and archaeological scientists in universities, museums, research laboratories, and field units. Each book includes a survey of current archaeological practice alongside essential reference material on contemporary techniques and methodology.


Books in the series

Clive Orton, Paul Tyers, and Alian Vince, POTTERY IN ARCHAEOLOGY

R. Lee Lyman, VERTEBRATE TAPHONOMY

Peter G. Dorrell, PHOTOGRAPHY IN ARCHAEOLOGY AND CONSERVATION, 2ND EDITION

A. G. Brown, ALLUVIAL GEOARCHAEOLOGY

Cheryl Claasen, SHELLS

Clive Orton, SAMPLING IN ARCHAEOLOGY

Steve Roskams, EXCAVATION

Simon Hillson, TEETH, 2ND EDITION

William Andrefsky, Jr., LITHICS, 2ND EDITION

James Conolly and Mark Lake, GEOGRAPHICAL INFORMATION SYSTEMS IN ARCHAEOLOGY

Andrew Chamberlain, DEMOGRAPHY IN ARCHAEOLOGY

A. M. Pollard, C. M. Batt, B. Stern, and S. M. M. Young, ANALYTICAL CHEMISTRY IN ARCHAEOLOGY





Zooarchaeology

SECOND EDITION

Elizabeth J. Reitz University of Georgia

Elizabeth S. Wing Florida Museum of Natural History





CAMBRIDGE UNIVERSITY PRESS
Cambridge, New York, Melbourne, Madrid, Cape Town, Singapore, São Paulo, Delhi

Cambridge University Press
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www.cambridge.org
Information on this title: www.cambridge.org/9780521857260

© Elizabeth J. Reitz and Elizabeth S. Wing 2008

This publication is in copyright. Subject to statutory exception
and to the provisions of relevant collective licensing agreements,
no reproduction of any part may take place without
the written permission of Cambridge University Press.

First published 2008

Printed in the United States of America

A catalog record for this publication is available from the British Library.

Library of Congress Cataloging in Publication Data

Reitz, Elizabeth Jean, 1946–
Zooarchaeology / Elizabeth J. Reitz, Elizabeth S. Wing. – 2nd ed.
   p. cm. – (Cambridge manuals in archaeology)
Includes bibliographical references and index.
ISBN 978-0-521-85726-0 (hardback) – ISBN 978-0-521-67393-8 (pbk.)
1. Animal remains (Archaeology) – Identification – Handbooks, manuals, etc.
I. Wing, Elizabeth S. II. Title. III. Series.
CC79.5.A5R45  2007
930.1–dc22      2007019451

ISBN 978-0-521-85726-0 hardback
ISBN 978-0-521-67393-8 paperback

Cambridge University Press has no responsibility for
the persistence or accuracy of URLs for external or
third-party Internet Web sites referred to in this publication
and does not guarantee that any content on such
Web sites is, or will remain, accurate or appropriate.





Dedicated to our families and colleagues in appreciation of their support and inspiration.





CONTENTS



List of Figures page ix
List of Tables xv
Preface to the Second Edition xix
Acknowledgments xxiii
1.   Zooarchaeology 1
2.   Zooarchaeological History and Theory 11
3.   Basic Biology 31
4.   Ecology 88
5.   Disposal of Faunal Remains and Sample Recovery 117
6.   Gathering Primary Data 153
7.   Secondary Data 182
8.   Humans as Predators: Subsistence Strategies and Other Uses of Animals 251
9.   Control of Animals Through Domestication 287
10.   Evidence for Past Environmental Conditions 316
11.   Conclusions 335
Appendix 1: Taxonomic List 353
Appendix 2: Anatomical Drawings 363
Appendix 3: Reference Collections, Management of Archaeofaunal Collections, Publication, and Curation 377
Appendix 4: Hypothetical Collection Data 396
Bibliography 409
Systematic Index 491
Topical Index 504




LIST OF FIGURES



  1.1  Locations mentioned in the text. page 3
  2.1  Section of an Aleutian shell heap. 15
  2.2  Fish hooks and the stages in their manufacture. 16
  2.3  Bone objects from Burial A-2, Uaxactun, Guatemala. 17
  2.4  Theodore E. White. 21
  2.5  Four behavioral strategies predicted by middle-range theory. 23
  2.6  Catchment area of an Arab village. 25
  3.1  Vertebral types. 42
  3.2  Dorsal views of (a) a shark (Carcharhinidae) and (b) a primitive bony fish (tarpon, Megalops atlanticus) vertebra. 43
  3.3  Three of the basic types of tooth attachments. 43
  3.4  Sea urchin test and feeding complex. 45
  3.5  Right lower first molar of a cow (Bos taurus). 47
  3.6  Different mammalian dental characteristics as seen in the upper dentition of (a) a herbivore, (b) a herbivore-omnivore, (c) a carnivore, and (d) a specialized carnivore. 48
  3.7  A porgy (Sparidae) dentary showing differentiation of teeth. 49
  3.8  A Florida stone crab (Menippe mercenaria) cheliped showing rounded cusps for crushing invertebrates. 50
  3.9  Left and right shrimp (Penaeus sp.) mandibles. 51
  3.10  A beaver (Castor canadensis) skull. 52
  3.11  Stoplight parrotfish (Sparisoma viride) jaws and pharyngeal grinding mills. 53
  3.12  A lateral view of the left dentary of a piranha (Serrasalmus sp.). 54
  3.13  Elements in the right front flipper of a whale (Cetacean). 57
  3.14  Elements in digitigrade (a and b) and unguligrade (c and d) left feet. 58
  3.15  Elements in the right wing of a bird. 59
  3.16  Gastroliths of a land crab (Gecarcinidae). 62
  3.17  Scatter plots showing the relationship between the total weight (Y) and the skeletal weight (X) of some terrestrial mammals. 67
  3.18  The tibia of a young cow (Bos taurus) showing the diaphysis and the unfused epiphyses. 71
  3.19  A southern quahog (Mercenaria campechiensis) showing the right valve, where it was sectioned (a), and a side view of the section (b). 74
  3.20  The first anal fin spine of a spadefish (Platax sp.) showing the hyperostosis of an interhaemal element. 75
  3.21  The projecting epiplastron of (a) a male gopher tortoise (Gopherus polyphemus) compared with (b) a female. 81
  3.22  Isotopic variation in nature. 85
  4.1  Diagrammatic relationship of nutritional requirements. 97
  4.2  Survivorship curves. 101
  4.3  Adjacent habitats in a tropical coastal setting with rocky splash zone, shallow inshore waters, fringing mangroves, and mountain headlands. 105
  4.4  The distribution of major terrestrial biomes with respect to mean annual temperature and mean annual precipitation. 107
  4.5  Decline in richness of mammalian species with increased degrees of latitude. 111
  4.6  Rarefaction curve of sample size (number of individuals) and species richness (number of genera) based on faunal samples from the West Indies. 114
  5.1  The possible pathway from a life assemblage to the archaeological assemblage with some of the major changes that can alter the assemblage. 119
  5.2  Kill hole in the umbo of an ark (Arcidae) made by a carnivorous marine snail (superfamily Muricacea). 122
  5.3  West Indian topsnail (Cittarium pica) showing modification of the shell by the resident land hermit crab (Coenobita clypeatus). 123
  5.4  The right scapula of a red deer (Cervus elaphus) with a healed fracture that pierced the blade. 125
  5.5  Kill hole in a lightning whelk (Busycon sinistrum) made by human foragers. 127
  5.6  Bone modifications from Mesolithic sites in Denmark: Impact marks. 128
  5.7  Bone modifications from Mesolithic sites in Denmark: Chop or hack marks. 129
  5.8  Bone modifications from Mesolithic sites in Denmark: Cut marks. 129
  5.9  Bone modifications from Mesolithic sites in Denmark: Scrape marks. 130
  5.10  Bone modification as a result of sawing. 131
  5.11  Gnaw marks made by (a) a rodent and (b) a dog. 137
  5.12  The size distribution of fish vertebral centra from the Kings Bay site on the Atlantic coast of Georgia, United States. 149
  6.1  Map of Spanish Florida showing the location of St. Augustine. 155
  6.2  Hypothetical Collection: Examples of data cards for white-tailed deer (Odocoileus virginianus). 159
  6.3  Hypothetical Collection: Drawing of white-tailed deer (Odocoileus virginianus) specimens recorded in the two samples presented in Figure 6.2. 162
  6.4  Hypothetical Collection: White-tailed deer (Odocoileus virginianus) skeleton summarizing specimens identified in the collection. 163
  6.5  Hypothetical Collection: An example of diagnostic zones for white-tailed deer (Odocoileus virginianus) radius specimens. 165
  6.6  Some typical modification characteristics. 169
  6.7  Linear enamel hypoplasia on the lingual surface of domestic pig (Sus domesticus) molars. 170
  6.8  Hypothetical Collection: Location of butchering marks on white-tailed deer (Odocoileus virginianus) specimens in the collection. 171
  6.9  Mandibular tooth wear stages for (a) pig (Sus domesticus), (b) cattle (Bos taurus), and (c) sheep/goat (Caprinae). 175
  6.10  Codes for caprine mandibular tooth wear stages. 177
  7.1  Hypothetical Collection: White-tailed deer (Odocoileus virginianus) measurements. 184
  7.2  Hypothetical Collection: Chicken (Gallus gallus) measurements. 185
  7.3  Hypothetical Collection: Steps to estimate Standard Length (SL) of red drum (Sciaenops ocellatus) from atlas width. 187
  7.4  Hypothetical Collection: Ratio diagram showing the size of white-tailed deer (Odocoileus virginianus) compared to the size of a modern standard. 190
  7.5  Hypothetical Collection: Using measurements to distinguish between Norway rats (Rattus norvegicus) and black rats (R. rattus). 191
  7.6  Hypothetical Collection: Histogram of white-tailed deer (Odocoileus virginianus) age groups. 197
  7.7  Hypothetical Collection: (a) Sea catfish (Ariidae) otolith length and (b) mullet (Mugil spp.) atlas width measurements. 198
  7.8  Characteristics of hard clams (Mercenaria mercenaria) based on increment counts and measurements of modern and archaeological (A.D. 1000–1500) clams from Kings Bay, Georgia. 199
  7.9  Age at death of Atlantic croaker (Micropogonias undulatus) from a sixteenth-century Contact Period deposit at the Fountain of Youth site near St. Augustine based on annuli in otoliths. 200
  7.10  Scatter diagrams of (a) modern Aberdeen Angus metacarpal bones and (b) bovine metacarpal bones from Troldebjerg, a Danish prehistoric settlement. 201
  7.11  Hypothetical Collection: Specimens identified as white-tailed deer (Odocoileus virginianus) shown anatomically using the Number of Identified Specimens (NISP). 218
  7.12  Hypothetical Collection: Histogram of white-tailed deer (Odocoileus virginianus) specimens. 219
  7.13  Hypothetical Collection: Illustrations of white-tailed deer (Odocoileus virginianus) radii and ulnae identified. 220
  7.14  Hypothetical Collection: Ratio diagram of white-tailed deer (Odocoileus virginianus) skeletal portions using NISP. 223
  7.15  Hypothetical Collection: Corrected frequencies for white-tailed deer (Odocoileus virginianus) using NISP. 225
  7.16  Hypothetical Collection: Food Utility Index (FUI) plotted against NISP percent for white-tailed deer (Odocoileus virginianus). 230
  7.17  Hypothetical Collection: Food Utility Index (FUI) plotted against %MAU for white-tailed deer (Odocoileus virginianus). 231
  7.18  Hypothetical Collection: Plot of volume density (VD) against NISP for white-tailed deer (Odocoileus virginianus). 233
  7.19  Hypothetical Collection: Estimates of white-tailed deer (Odocoileus virginianus) dietary contribution based on concepts of individuals using total weight or measured dimensions. 235
  7.20  Hypothetical Collection: Estimates of white-tailed deer (Odocoileus virginianus) dietary contribution based on archaeological specimen weight. 236
  7.21  Hypothetical Collection: Comparing the results of methods that estimate individual weights and sample biomass for (a) white-tailed deer (Odocoileus virginianus) and (b) bony fishes. 241
  7.22  Hypothetical Collection: Butchering marks for white-tailed deer (Odocoileus virginianus) shown on a skeleton. 244
  7.23  Hypothetical Collection: Histogram comparing burned and unburned specimens for mammals, birds, turtles, and fishes. 245
  7.24  Calculating diversity and equitability using the Shannon–Weaver and Sheldon formulae. 247
  8.1  A hypothetical example of regional settlement patterns and socioeconomic changes from the Middle Archaic to the Hohokam Period in the southwestern United States. 258
  8.2  Map of the Lower Pecos Region, the Chihuahuan Desert, United States and Mexico, illustrating the seasonal round hypothesized by Shafer. 259
  8.3  Seasons of collecting for hard clams (Mercenaria mercenaria) based on characteristics of narrow translucent (T) growth increments and broader opaque (O) increments. 263
  8.4  Instruments: (a) Surprise Valley Paiute digging stick, southwestern United States; (b) Klamath digging stick, Northwest coast; (c) Owens Valley Paiute seed beater, southwestern United States; and (d) Twana herring rake, Northwest coast. 269
  8.5  Simple and complex weapons: (a) Simple weapon, a Tanala leister, Madagascar; (b) complex weapon, an Ingura dugong harpoon dart, northern Australia; and (c) complex weapon, an Iglulik bird dart and throwing-board, northern Canada. 269
  8.6  Tended facilities: (a) Tanala cone-shaped fish scoop, Madagascar; (b) Tanala dip (scoop) net, Madagascar; (c) Klamath composite fish hook, Northwest coast; (d) Tlingit halibut hook, Northwest coast; (e) Klamath fish gorge, Northwest coast; and (f) Twana salmon weir and dip net platform, with profile of weir, Northwest coast. 271
  8.7  Untended facilities: (a) Simple snare, Pukapukan baited coconut shell bird snare, Cook Islands; (b) complex trap, Tanaina torque trap, Alaska; (c) Tanala baited eel trap, Madagascar; and (d) Ingalik whitefish net set beneath river ice, Alaska. 273
  8.8  Crocodilian image from an “Alligator Ware” vessel, Chiriqui province, Panama, painted between A.D. 1100 and 1520. 283
  9.1  Shortened snout and high forehead traits found in many domestic animals. 305
  9.2  Llama pack train in the Andean highlands. 308
  9.3  Pottery model of a llama sitting down but loaded with a woven bag on its back and identification notches in its ear. 309
  10.1  Von Bertalanffy growth curve fit to back-calculated size at age for Atlantic croaker (Micropogonias undulatus). 327
  A2.1  Directional terms for vertebrates using a pig (Sus domesticus). 363
  A2.2  Dog (Canis familiaris) skeleton with some elements labeled. 364
  A2.3  Bird skeleton with some elements labeled. 364
  A2.4  The cervical vertebrae of the snapping turtle (Chelydra serpentina). 365
  A2.5  Turtle (a) carapace and (b) plastron with some elements labeled. 365
  A2.6  Skull of West African broad-fronted crocodile (Osteolaemus tetraspis). 366
  A2.7  Skull of iguana (Ctenosaura pectinata). 367
  A2.8  Skull of African spitting cobra (Naja nigricollis). 368
  A2.9  Python (Python sp.) and iguana (Iguana sp.) vertebrae. 369
  A2.10  Frog (Ranidae) skeleton with some elements labeled. 370
  A2.11  The cranium of a striped bass (Morone [=Roccus] saxatilis) with some elements labeled. 371
  A2.12  Lateral facial elements and appendicular skeleton of a striped bass (Morone [=Roccus] saxatilis). 372
  A2.13  The axial skeleton of a striped bass (Morone [=Roccus] saxatilis). 372
  A2.14  A West Indian fuzzy chiton (Acanthopleura granulata). 373
  A2.15  Internal and external features of a spiral gastropod shell (Gastropoda). 373
  A2.16  Dorsal view of a clam (Bivalvia). 374
  A2.17  Inside view of the left valve of a clam (Bivalvia). 374
  A2.18  Crustacean external morphology of a crayfish (class Malacostraca, infraorder Astacidae). 375
  A2.19  General anatomy of a swimming crab (class Malacostraca, order Decapoda, Portunidae). 375
  A2.20  Left mandible of a land crab (Gecarcinidae). 376
  A3.1  Some standard measurements in birds (Aves) and mammals (Mammalia). 380
  A3.2  Some standard measurements in turtles (Testudines), lizards (Lacertilia), and snakes (Serpentes). 381
  A3.3  Some standard measurements in fish (Actinopterygii), salamanders (Caudata), and toads and frogs (Anura). 382
  A3.4  Some standard measurements for eastern oysters (Crassostrea virginica), hard clams (Mercenaria spp.), and knobbed whelk (Busycon carica). 383




LIST OF TABLES



  3.1  The higher classification of the stoplight parrotfish (Sparisoma viride), with a brief list of the characteristics of each category. page 35
  3.2  Classes most commonly found in archaeological contexts and characteristics of their most common remains. 40
  3.3  The relative percentage of organic and inorganic constituents of various hard tissues. 41
  3.4  Some allometric regression formulae. 68
  3.5  Age in months when epiphyseal fusion of different skeletal elements may be completed in some mammals. 72
  4.1  Life table constructed for female age distribution among nomadic cattle of east Africa. 102
  4.2  Net annual primary productivity and standing crop biomass estimates for contrasting communities of the world. 108
  4.3  Example of the calculation of species diversity in (a) a sample in which four species are equally represented compared with (b) a sample in which one of the four species predominates. 112
  4.4  Calculation of the percentage similarity among the average number of domestic animals kept by peasants with small, medium, and large herds in the Laguna Blanca Reserve, Argentina. 115
  6.1  Hypothetical Collection: Vertebrate summary showing the impact of screen size on the Number of Identified Specimens (NISP). 156
  6.2  Hypothetical Collection: Horn core attributes for cattle (Bos taurus). 179
  7.1  Hypothetical Collection: White-tailed deer (Odocoileus virginianus) measurements, range, mean, standard deviation (SD), and sample size (N ); measurements in mm. 188
  7.2  Hypothetical Collection: Data used to construct a ratio diagram for white-tailed deer (Odocoileus virginianus); measurements in mm. 189
  7.3  Hypothetical Collection: White-tailed deer (Odocoileus virginianus) NISP subdivided into age categories. 195
  7.4  Hypothetical Collection: Skeletal groups for fishes, NISP. 205
  7.5  Hypothetical Collection: NISP of white-tailed deer (Odocoileus virginianus), diamondback terrapin (Malaclemys terrapin), gopher tortoise (Gopherus polyphemus), and pit viper (Viperidae) in five archaeological contexts. 209
  7.6  Hypothetical Collection: White-tailed deer (Odocoileus virginianus) MNI estimates subdivided into age categories and archaeological contexts. 210
  7.7  Hypothetical Collection: Summary of NISP, MNI, specimen weight, and sample biomass data. 211
  7.8  Hypothetical Collection: Calculation of percentage similarity among some vertebrates from a sixteenth-century Spanish site in St. Augustine; an eighteenth-century African site, Fort Mose; and the Hypothetical Collection, using MNI. 214
  7.9  Hypothetical Collection: Summary of some mammalian specimens by anatomical regions, NISP. 217
  7.10  Hypothetical Collection: Observed/expected ratio for white-tailed deer (Odocoileus virginianus); expected based on NISP. 221
  7.11  Hypothetical Collection: Percentage survival for white-tailed deer (Odocoileus virginianus) based on the number of each element expected for eleven individuals. 222
  7.12  Hypothetical Collection: Meat utility indices and bone mineral densities for white-tailed deer (Odocoileus virginianus). 228
  7.13  Hypothetical Collection: Rank of white-tailed deer (Odocoileus virginianus) elements based on NISP percentage, corrected frequency, MNE percentage, and %MAU. 229
  8.1  Characteristics of ritual animal use, broadly defined. 285
  9.1  Major domestic animals, their presumed wild ancestors, region(s) of domestication, and approximate date of first domestication. 291
  11.1  Some correlations between primary and secondary data and related concepts. 336
  A1.1  Taxonomic list that includes all animals mentioned in the text. 353
  A3.1  Curatorial information and primary data included on labels and data cards. 388
  A3.2  Checklist for zooarchaeology reports. 391
  A4.1  Hypothetical Collection: Species list. 396
  A4.2  Hypothetical Collection: Number of specimens with modifications. 401
  A4.3  Hypothetical Collection: Specimen distribution worksheet for white-tailed deer (Odocoileus virginianus). 402
  A4.4  Hypothetical Collection: Fusion worksheet for white-tailed deer (Odocoileus virginianus). 405
  A4.5  Measurements from the Hypothetical Collection and other collections used in examples. 406




PREFACE TO THE SECOND EDITION



When we were asked to prepare a second edition to Zooarchaeology, we anticipated that this would be relatively easy. We proposed to update the literature and work on sections that we or our colleagues found did not “work” in practice. We quickly realized, however, the truth of the statement that zooarchaeology is a dynamic field. We were surprised to find a few major changes in the traditional approaches in the field over the past 10 years and significant advances in archaeogenetic, isotopic, and incremental growth applications. A shift in research emphasis also has occurred. Whereas in 1999 many zooarchaeologists focused on biological and anthropological interpretations pertaining to economies and the history of animal domestication, today publications on environmental change, environmental reconstruction, and applied zooarchaeology constitute a large percentage of the literature. Advances in geochemical applications make it possible to develop holistic perspectives on the human–environment relationship, dissolving problematic distinctions among anthropology, archaeology, ecology, geology, human biology, and zoology. At the same time, after many years of functional interpretations, structural explanations have assumed a larger place in the literature. One of the most gratifying discoveries is the increase in important zooarchaeological studies published in peer-reviewed, international journals by scholars from beyond Europe and North America. This more broadly inclusive community of scholars is a good sign that zooarchaeology continues to be strongly international.

   Thus, in preparing this second edition, we made major changes in sections in which the greatest advances have been made in the past decade. Chapters 3, 9, and 10 are substantially rewritten to incorporate new information and research trajectories. Chapter 8 required significant, though less extensive, modifications. We have included literature from 1999 through 2006, as well as inadvertent omissions from before 1999. These new references reflect the directions that the field of zooarchaeology has taken over the past decade. We believe these directions represent the future of the field. We repeat, however, our admonition from the first edition that theoretical interpretations are no better than the methods used to develop supporting data. It is as necessary to be well-grounded in the basics as it is to be guided by good theory.

   There are some changes we did not make. As with the first edition, this edition is not intended to replace the many excellent biological references; works focused on single organisms or groups of organisms; methodological descriptions and reviews developed out of specific research needs; or regional archaeofaunal syntheses and theoretical treatments. The focus of the volume continues to be on topics of broadly global applications pertaining to major research trajectories. We do not advocate for or against methods or provide detailed descriptions of specific methods or outcomes. Students should recognize the importance of developing their own research designs, which will be implemented using appropriate methods obtained through their own review of the literature.



PREFACE TO THE FIRST EDITION

This volume is directed to all those interested in the recovery, identification, and ana-lysis of animal remains from archaeological sites. Our intent is to review standard zoo-archaeological methods and to suggest the circumstances under which they may be most successfully applied. Because we believe that a background in both anthropology and biology is important for a balanced approach to zooarchaeology, both relevant anthropological and biological information are reviewed. The exchange among archaeological, biological, ethnographic, and paleontological research is the important defining characteristic of the study of animal remains that links the following pages. The development of zooarchaeology owes much to an awareness of the importance both of ecological relationships on human behavior and of the human impact on the planet. Despite its diverse, interdisciplinary nature, zooarchaeology has three common research themes: (1) methodology; (2) continuity and change in human societies; and (3) biological relationships. These are the primary topics explored in this volume.

   The animals emphasized include macrofaunal as well as some microfaunal organisms. The term “macrofauna” refers to large vertebrates and invertebrates. All vertebrate classes are included. These are mammals (Mammalia), birds (Aves), reptiles (Reptilia), amphibians (Amphibia), cartilaginous fishes (Chondrichthyes), and bony fishes (Actinopterygii). Invertebrates include primarily molluscs (Mollusca) and crustaceans (Crustacea). The term “microfauna” may refer to small members of these same classes, such as anchovies, or to small organisms, such as land snails or insects. The tissues reviewed include skeletal bone and teeth, mollusc shell, and exoskeleton (such as crab shell). Egg shell and keratinized tissue, such as hair, skin, and feathers, are not stressed here.

   Our emphasis is on animals whose remains inform us about aspects of relationships between humans and their natural and social environments, especially site-formation processes, subsistence strategies, and paleoenvironments. Among these animals, those that offer food, shelter, transport, fuel, tools, ornaments, clothing, and social identity receive particular attention. We also explore material culture related to the procurement and husbandry of animals. Examples are primarily those illustrating modern human (Homo sapiens sapiens) uses of these animals. The time period is from the Pleistocene into the twenty-first century A.D.

   The geographic range is global. Although examples are drawn from many parts of the world, we make no effort to provide regional surveys of zooarchaeological developments. Smith’s (1995) review of the emergence of agriculture throughout the world provides regional surveys of both plant and animal data. His volume is a good place to obtain an overview of zooarchaeological knowledge in the context of broader archaeological research. Our intention is to review anthropological, biological, and ecological aspects of zooarchaeology from the wide variety of geographical settings in which zooarchaeology is practiced and to summarize, broadly, the diverse ways in which humans and animals interact.

   The volume is organized in much the same way as a faunal study might be. Familiarity with the history of zooarchaeology and current research topics provides the intellectual background a zooarchaeologist should bring to the study of a specific faunal assemblage (Chapter 2). It is also important to be familiar with biological (Chapter 3) and ecological principles (Chapter 4) basic to the discipline. In Chapters 5, 6, and 7, three sources of bias in a faunal assemblage are reviewed, beginning with taphonomy and excavation procedures. Chapters 6 and 7 present some of the most basic zooarchaeological methods, using an archaeofaunal Hypothetical Collection to illustrate fundamental methods for collecting primary and secondary data. In the remaining chapters, animal remains are interpreted in terms of subsistence strategies (Chapter 8), domestication (Chapter 9), and human interactions with the environment (Chapter 10). The final chapter (Chapter 11) draws these threads together and considers future directions in the field.

   This volume is not intended to replace the many excellent biological references available; works focused on single organisms or groups of organisms; methodological descriptions and reviews; regional archaeofaunal syntheses; or theoretical treatments. Extensive references are offered for each topic covered in the following pages. We urge readers to use these as guides to more detailed treatments of each subject. We hope to excite students to pursue their own interests in this diverse field so that they may share, with us, many hours of stimulating puzzlement.





ACKNOWLEDGMENTS



It is with deep gratitude that we acknowledge the contributions of the many people who helped to bring the first and second editions of this volume to completion. In particular, we appreciate the willingness of Graeme Barker, Laszlo Bartosiewicz, Don R. Brothwell, Anneke T. Clason, Dena F. Dincauze, Norman Herz, Stephen A. Kowalewski, Clark Spencer Larsen, William H. Marquardt, Arturo Morales Muñiz, Lynette Norr, Barnett Pavao-Zuckerman, Ann B. Stahl, and an anonymous reviewer to read all or portions of the manuscript. Richard G. Cooke, Simon J. M. Davis, Annie Grant, Laura Kozuch, Robert Newman, Nanna Noe-Nygaard, Wendell H. Oswalt, Paul W. Parmalee, Sebastian Payne, Irvy R. Quitmyer, and Melinda A. Zeder were particularly generous in helping to prepare the illustrations. Sarah M. Colley, Greg Cunningham, Elizabeth McGhee, Katherine E. Quitmyer, Dawn Reid, Donna Ruhl, Jaap Schelvis, Pamela Soltis, David W. Steadman, and Stephen R. Wing also provided invaluable assistance at critical times. Special thanks are extended to Max Reitz, Sylvia J. Scudder, Irvy R. Quitmyer, and the students in our zooarchaeology classes. We may not have followed all of the insightful suggestions offered, but we are grateful for the comments nonetheless. The volume is enhanced by the artistic contributions of Virginia Carter Steadman, Paloma Ibarra, Tina Mulka, Tara Odorizzi, Daniel C. Weinand, Molly Wing-Berman, and Wendy Zomlefer. The cover illustration was prepared by Molly Wing-Berman. Most of the graphics not attributed to these artists were prepared by Gisela Weis-Gresham, who once again demonstrates the value of good illustrations. We are grateful to the many individuals and presses who granted permission to use work previously published. We also appreciate the advice and support of Beatrice Rehl, Peggy M. Rote, Jessica Kuper, Frances Brown, and the Cambridge University Press staff. Through the generosity of Kitty Emery, the Florida Museum of Natural History provided essential support during the final preparation of the second edition. Finally, we thank the Florida Museum of Natural History, the Georgia Museum of Natural History, colleagues (among whom we number our students), and families for their patience and cooperation as we attempted to squeeze the field of zooarchaeology into a single volume.





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